Ask any SEO expert this question and most of them will answer the same thing that links from non-relevant sites are not only valueless but also have a negative effect your site's ranking on search engines.
In simple words, these links will pass PageRank but will have little or no effect on the SERPs. If you add too many of these on your site then it will have a negative effect on SERPs. It is important to understand that PR (PageRank) matters only when all other things are equal. Instead, if you will be better off getting back links from a relevant with targeted anchor text rather than getting back links from an irrelevant sites.
Therefore, make sure to link sites that has relevant content irrespective of its PR. So you see, it's all about getting valuable links to make your website available to users of other related websites. You'll find many softwares which helps in managing 'Linking Programs'. Some of this software is 'automated' and allows links from anyone who is willing to fill in a form and post a link to their website first.
If you're wondering how high PR if help your SE rankings, then you'll be surprised to know these programs have been in the very least, achieving nothing for the SE rankings by collecting so many irrelevant links. Hence, it's possible that in so doing, they have been adding a burden for your website and may well have been penalized for it.
However, do not misguided by the fact that link popularity alone is considered in the SE ranking algorithms. Instead, it is important to note that links from a non-related site will still pass PR, but will have little or no effect on the SERPs.
Therefore, understand one thing clearly that linking for the purpose of improving SE rankings, only works if it's done properly. This means, to exclude irrelevant links from your site and webpages.
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